Wednesday, July 19, 2023

Fossil prep updates: beluga and fur seal specimens from the Purisima Formation of California

We had some great luck while on our May field trips in coastal California, as reviewed in my last couple of posts. Sarah and I have added a few more scientifically significant specimens to the faunal list for this locality. The fossil vertebrate assemblage from the Purisima Formation is certainly not the most numerous in terms of specimens, but is rapidly becoming the most completely published. In 2013 I published a hefty monograph on the marine mammal assemblage from the locality, following an earlier paper reporting all of the sharks, bony fish, and sea birds - and a followup that same year reporting the world's youngest well-dated bony-toothed bird fossil (Pelagornis). Localities like this are quite critical since they are the youngest diverse assemblages of marine vertebrate fossils from our coast: Pleistocene marine vertebrate assemblages are poorly known from the west coast and typically non-comparable in terms of diversity, given that they are typically 'singleton' records and not evenly sampled for both "lower" and "higher" vertebrates. The new records I'll be discussing in this post are marine mammals, though we've made some interesting new non-mammal additions which I'll cover in a later post. 


 This photo is unintentionally hilarious - I was *so* happy by this fossil! Sarah made no mention of the fact that you cannot tell what the hell I am actually holding, it just looks like a shitty little chunk of sandstone. Didn't matter, I was so happy, and this is far funnier than if it was this spectacular field glory photo.

The first noteworthy specimen is a partial mandible with teeth of a tiny little fur seal. Pinniped remains are quite rare, outnumbered by cetacean fossils in the Purisima Formation by at least 10 to 1 or 20 to 1 (they are over-represented in UCMP collections, but even then still outnumbered by cetaceans by 5 to 1). Most pinniped fossils from the Purisima Formation consist of isolated limb elements and teeth; skulls, mandibles, and fragments thereof are considerably more rare. Mandibles are frequently diagnostic within pinnipeds, and so I had always been on the look out for one at this locality. I had previously collected a partial radius of a dusignathine walrus (Dusignathus sp.) and a very nice calcaneum and upper third incisor from a small fur seal - along with specimens collected by Larry Oliveira (such as a nearly complete but very small humerus), I tentatively identified this fur seal as the extinct Callorhinus sp., cf. C. gilmorei.


The holotype (female) mandible of Callorhinus gilmorei from the Pliocene San Diego Formation.

Referred mandibles of Pithanotaria starri from the upper Miocene Monterey Formation of Orange County, CA. From Velez-Juarbe (2017: PeerJ 5:e3022) .

Callorhinus gilmorei just so happens to have a pretty distinctive mandible, even if the teeth aren't terribly distinctive. It is one of the few fossil otariids where only a couple of the premolars are single rooted, and the rest are double rooted; the original specimen is a fragmentary but nonetheless well-preserved female skeleton from the coeval San Diego Formation. Other mandibles show a bit of variation, and at least one mandible has all double-rooted teeth. You see, pinnipeds evolved from terrestrial mammals that had double rooted premolars and triple rooted molars, and this condition was retained in the earliest pinnipeds like Enaliarctos. Early fur seals and sea lions (Otariidae) from the middle and late Miocene, such as Eotaria, Pithanotaria, and Thalassoleon, mostly have double rooted teeth - though all modern fur seals and sea lions have single rooted teeth. 


 Partially prepared mandible fragment with in situ premolars collected back in May.

This new mandible I collected in May ended up having all double-rooted teeth, which alone would not prevent this specimen from representing Callorhinus gilmorei. However, unlike this Pliocene species, this specimen has relatively smooth enamel that lacks accessory cusps and completely lacks lingual cingulum: cingulum is a little ridge on the base of the tooth crown. Most modern fur seals and sea lions have cingulum along the tongue side (lingual = tongue side) and only a few sea lions have labial cingulum. This specimen, however, has neither labial nor lingual cingulum, and the one complete tooth is also a bit low crowned. These features make this specimen resemble Pithanotaria - which, so far, is only known from the late Miocene (Tortonian, ~9-11 Ma). This specimen is probably, but uncertainly early Pliocene in age.

Another such specimen I found ended up being the anterior (chin) end of a mandible of a relatively large fur seal, collected from a bonebed dating to the early Pliocene - and a bit further up-section. After a bit of preparation, it ended up having single-rooted first and second premolars, identifying the specimen as Callorhinus gilmorei, and likely confirming my earlier identification of specimens of Callorhinus sp., cf. C. gilmorei from these same layers.


 The beautiful beluga periotic I collected in May! My very first monodontid periotic. I've only been looking for one for 15 years. Even on the east coast where they're more common!


 The new associated periotic and bulla of a small monodontid, collected under permit from the Purisima Formation.

One of the other really interesting specimens that I collected was an associated periotic and tympanic bulla of a large odontocete - which I was immediately able to identify to the family level in the field, owing to its distinctive anatomy: it is from a white whale, family Monodontidae - closely related to modern belugas and narwhals (and probably more the former). If you're unfamiliar with the fossil record of monodontids - and, surprisingly, I've never ever covered them here before (I should change that) - they parallel the walrus fossil record in a few surprising ways. First, they show up during the late Miocene, and have a similar distribution in the temperate and even subtropical North Pacific and North Atlantic. Aside from a couple of fragmentary finds of fossil narwhals, most are attributable instead to "delphinapterines" - tuskless monodontids more closely resembling belugas (Delphinapterus) than the narwhal (Monodon). These include species like Denebola brachycephala from the latest Miocene of Baja California, Bohaskaia monodontoides from the early Pliocene of North Carolina, and the more recently named Haborodelphis japonicus from the early Pliocene of Japan. There is additionally an unnamed fragmentary monodontid from the Pliocene of Belgium briefly described by my colleague Olivier Lambert in 2007, and several unnamed species of monodontids in the late Miocene-Pliocene fossil assemblages of California known from partial to complete skulls, possibly representing two or more genera.

 

The fractured skull (left) and periotic (right) of the small pilot whale-convergent beluga Denebola brachycephala from the upper Miocene Almejas Formation of Baja California, Mexico. From Barnes (1984: PaleoBios 42:1-46).

The small beluga Haborodelphis japonicus from the lower Pliocene Embetsu Formation of Hokkaido, Japan. From Ichishima et al. (2019: Papers in Palaeontology 5: 323-342).
 


 The periotic and tympanic bulla of
Haborodelphis japonicus. From Ichishima et al. (2019: Papers in Palaeontology 5: 323-342).

Out of these three named fossil species, only Denebola and Haborodelphis have periotics preserved that can be directly compared with my new specimen. One unnamed monodontid from a different locality in the Purisima Formation a few miles away has periotics that are a bit dorsoventrally thicker than my new one. The new one seems to have a proportionally larger anterior process than Denebola, and more closely resembles Haborodelphis. The tympanic bulla, on the other hand, is perhaps a bit more informative: the thickened part of the bulla, the involucrum, is absurdly thick anteriorly in Denebola and bears a long anterior tip; the new specimen looks quite a bit more like Haborodelphis and many other delphinoids. The bulla of the new specimen is only 3/4 the size of Denebola, yet the periotic is somehow larger. These are cursory observations, but I'm inclined to identify the specimen as cf. Haborodelphis for the time being. Finding fossils of marine mammals in California belonging to species originally discovered in Japan should not be surprising: Desmostylus and Paleoparadoxia (now Neoparadoxia), Callorhinus and Eumetopias, Herpetocetus, Hydrodamalis, and many others have been found in both places.

Monday, June 12, 2023

Fieldwork snapshots: juvenile Herpetocetus mandible and more in the Purisima Formation

It's been a very rough couple of weeks for Sarah and I, so I apologize for being MIA. I'll explain later. But, for the time being, we're back to regular posting.

 Sarah below some rather imposing cliffs.

After our marathon day out in Half Moon Bay I realized there were a couple of localities we didn't get to return to, and one specimen in particular that remained uncollected. For some reason I thought that I had first spotted it in fall 2019 and I thought "Oh, I'll come back and collect it in the summer when I have a permit" and then Covid lockdown happened - it did dash my plans for summer fieldwork and I didn't bother applying for a new permit at the time. Going back through my photos, I realized I had actually spotted this fossil in December 2016! I then recalled revisiting the locality several times and failing to re-locate the specimen: a tiny little baleen whale mandible that was perhaps only 4 cm in height, which I figured must have been from a juvenile Herpetocetus - my favorite little dwarf baleen whale. The mandible was eroding out on a bit of a flat bench, so it was at less danger of being washed away - but it was frequently covered with loose bits of rock and dust that fell off the cliff face. On our prior day out there with Jorge Vazquez, Marsha Lidzbarski and Wayne Thompson, I had re-located the specimen - but it was quite a bit further up the cliff face than I had remembered, and I immediately recalled searching the wrong area once I found it. At least a few inches of the specimen had been eroded away forever - a bit of a bummer, but not bad considering I first spotted it seven years prior! I knew from the direction that the mental foramina opened on the side of the mandible that the diagnostic posterior end remained inside the cliff. These foramina (holes) are located on the tooth-bearing part of a regular mammal mandible, but in baleen whales, this part is rib-shaped and of course toothless - and they have more mental foramina than a land mammal, and they have long, anteriorly opening sulci. So - you can immediately tell which direction a mandible is going if you see just a little bit of it. If it's going straight into a cliff, it's a lot more difficult, nigh impossible, to tell.


 The juvenile Herpetocetus mandible as I first saw it in December 2016, partly washed 
off by winter storms.

Sarah and I hiked down the beach and passed a mollusk-bearing bed at the base of the cliff. Sarah froze her ass off the last time we had been out, and I promised her we'd keep today's field trip to about three hours. So, I suggested that she try and collect some mollusks while I dug out the Herpetocetus mandible. I climbed up the ledge and started carving away: the rock was very soft, which made me worry about the integrity of the bone. Earlier collections from this stratum resulted in rather crumbly bone. When I started, I also noticed that the mandible was just beginning to flare out into a ridge - I was pretty sure the coronoid process* of the mandible was present, and complete at that. After a little more digging, I identified the mandibular condyle* and knew that the entire posterior end was fortunately intact. 

*The coronoid process is what the temporalis muscle inserts into on the mandible. The condyle is the jaw joint.


 The mandible in May 2023, nearly seven years later - at least a few inches had eroded away.
 

 Sarah with her well-preserved scallop, possibly
Patinopecten healyi.

I waved to Sarah, about a hundred yards away, motioning that I was almost done. She returned to the ledge and pulled out a very well-preserved scallop shell that she had expertly collected: I had seen that individual scallop sticking out of the cliff, but left it, since I assumed I would break it. She had collected it in once piece, and even the little ears (auricles) were preserved near the umbo. I was impressed and congratulated her on the find! Such scallops are often crucial index fossils for Neogene fossil localities - though in this case, as covered in the prior post, this section of the Purisima Formation is soon to become the most tightly geochronologically constrained marine section for the Pliocene anywhere on the west coast, to a far greater degree than invertebrates can help.


Excavating the tiny whale jaw from the ledge. Despite the way this photo looks, I'm actually about 20 feet (~7 meters) above the beach here.


Some mollusks from just above the mandible - mostly
Anadara trilineata, the common late Miocene-Pliocene ark clam from the west coast.

After a few more minutes of carving a trench into the soft sandstone and outlining a brick-shaped (and sized) block containing the mandible, I stuck my wood chisel horizontally and began to undercut the block with a few gentle taps. The block popped loose rather easily - and then in half, unfortunately. However, it was a very clean fracture, so I wrapped up each block separately with a few sheets of paper towel.

Teichichnus pescaderoensis burrows in the Purisima Formation.
 
After everything was bagged up, we headed down to the last locality. In this section, the Purisima Formation is dominated by the trace fossil Teichichnus pescaderoensis, a rather large trace fossil indicating relatively deep deposition on the outer shelf or slope, but as shallow as middle shelf. The trace itself looks like other Teichichnus but is enormous - up to 1.2 meters wide according to the original paper, but I've seen individual traces up to 1.5 meters. The original authors considered the trace too large for an annelid worm, and suggested that some kind of infaunal deposit feeding crustacean would dig down, form a horizontal tunnel or gallery, and then excavate sediment from the roof and the gallery would migrate upwards.
 
 
Some more Teichichnus pescaderoensis burrows in a fallen block of the Purisima - rotated to match stratigraphic up (the burrows themselves are good geopetals - they're always concave up).
 
There are no macroinvertebrates in this section, and vertebrates are rare, but there's at least one good bonebed with quite a bit of material in it. I've collected a smattering of decent specimens there over the years.
 

Here's a pretty messed up lumbar vertebra of a medium sized cetacean, likely a small baleen whale, that's all fractured and faulted. Specimens in this part of the unit are frequently fairly faulted apart.


I actually found this one while I was taking a leak - I get bored, and always look around. The habit has paid off in the past and I've made some decent finds - but this one is quite nice. It's an upper lateral tooth of a cowshark, probably Hexanchus griseus.


Sunday, May 21, 2023

Fieldwork snapshots: fur seals, belugas, and ash deposits in the Purisima Formation, May 2023

Earlier this week Sarah and I flew home to the SF Bay Area to visit family and find some fossils. We've done several days of chilly fieldwork along the beautiful but cold and foggy California coast and are now enjoying a weekend day indoors with my folks (though we'll be visiting one last locality for an hour or so this afternoon). We're lightly sunburned, sore, and surprisingly bruised after lots of up-and-down hiking over rock exposures and boulder falls. Fortunately, neither of us has come into contact with poison oak, been slammed by a wave, or sustained any moderate injuries (though I was about a second away from losing my big toenail, and Sarah did slip at one point - see below).

I won't reveal the exact location as it is not in Santa Cruz, but it is protected and I have received a permit to collect here, with specimens destined for UCMP collections - and a permit is required. It's admittedly a reasonably remote site, perhaps my favorite locality, and one I've been visiting now for nearly 20 years - and I'm on my third permit for the locality. I had a bit of luck here in December, and I hoped to have similar luck this week.

 Jorge Vazquez explaining their findings and dating of the Putah Tuff-correlative ash bed forming this ledge in the Purisima Formation.

We met USGS/Stanford geologists Jorge Vazquez and Marsha Lidzbarski out along the coast along with our buddy Wayne Thompson (retired teacher and CEO of Pacific Paleontology, LLC) for a long day of fieldwork. Jorge and Marsha have been working on dating ash beds in the Purisima. I was familiar with Andre Sarna-Wojcicki's efforts to chemically fingerprint ash deposits, linking them to the volcanic deposits, lava flows, and tuff beds near the volcanic vent where the rocks were dated better - this is a form of geochemical correlation of ashes without directly dating the ash beds themselves. Only two ash beds out at this section had previously been dated - one correlated with the ~2.5 Ma Ishi Tuff in the southernmost cascades and another correlated with the 3.35 Putah Tuff from the Sonoma Volcanics. Rather than just being chemically fingerprinted, Jorge, Marsha, and others were directly dating the ash layers within the Purisima section.

Incredible high-contrast fossil burrows infilled with volcanic ash from a thick ash bed correlated with the Ishi Tuff.

About two years ago I was invited to review a paper reporting ash dates directly from this section of the Purisima Formation I had labored for years over - these geologists were unaware of the paleontological significance of the section (e.g. documenting the chronology and faunal changes over the past three million years) and I was completely unaware that anyone else cared about this locality! I had a robust list of suggested changes, which ironically led Jorge, Marsha, and others to seek out more and more ash beds to sample. We've been intermittently chatting over the past few months and decided to meet up - go out to the cliffs, and compare notes. I was absolutely floored by how many additional ash beds there were out there, and many of them have since been sampled! I'm not used to geological study on the Pacific coast being so vigorous and rapid... usually west coast geologists seem to work on 'geologic time'. Some of my vertebrate localities now have incredible dates constraining their ages to intervals as brief as 200,000 years!

A small pinniped finger bone, likely a metapodial or phalanx.

We had two main goals: 1) check out as many different vertebrate-bearing horizons as possible and recover whatever we could and 2) walk though as much of the section as possible and exchange notes. Wayne's job as a permit co-signer was to become familiarized with the best localities so that he could sample them intermittently while I'm on the east coast. I expect to extend this permit a year into mid-late 2024, and can only afford to make it out here a couple times per year.

Every time I visit the Balaenoptera bertae type locality I have to stop by and pose with it. It's my first named species! And I found it, excavated, prepared and studied it. It doesn't get much better than that.

We started off in the uppermost part of the section, and showed everyone the type locality of Balaenoptera bertae - owing to extreme winter erosion from January through March, caused by back to back atmospheric river storms, the sand level was quite a bit lower than typical and the hole, once about a meter above the beach when I discovered and collected the skull in August 2005, was now about five meters above the beach! We walked a ways along the beach, hopeful to survey a large section of cliffs that are typically easily accessible at high tide - but got turned back as the waves were crashing down upon the only real 'choke point' here, a spur of rock that stuck out about 200' from the main line of the cliffs into the Pacific. On our side of the spur, I relocated two baleen whale mandibles I hadn't seen in years. The first was a humpback whale sized mandible I first spotted in 2005, which had the mandibular condyle protruding from the cliff. This mandible was sticking straight into the cliff and would have required a hole approximately three meters deep to excavate - far too much effort for a single mandible. Then, when I returned for more fieldwork in 2010 on my second permit, I couldn't relocate the mandible. I spotted it for the first time in 18 years because the cliffs had been cleaned off - the mandible had fractured flush with the cliff face, and the cliffs here are frequently quite covered with dust.

 

Sarah and the re-discovered baleen whale mandible. There's about three more meters of bone sticking straight into the cliff.

The second was a much smaller mandible which I spotted in December 2016, but as I didn't have a permit, I took some photos and intended to return with a permit sometime and collect it. It was the posterior half of a mandible of a small baleen whale, very likely to be my favorite dwarf baleen whale Herpetocetus. When I tried finding it in December 2022, I couldn't relocate it. I had a strong feeling that I would easily find it on this trip, so I ascended the bluff - and couldn't find it. Weird! Maybe it had eroded away. I walked further, looking for other fossils - and then spotted it, about 10 meters further up the bluff than I had remembered. Quite a bit of the mandible had eroded away, but fortunately the diagnostic posterior end, including the coronoid process, angular process, and mandibular condyle were intact - and their morphology confirmed my identification of the specimen as Herpetocetus. If I hadn't scored the rest of this specimen it would have almost certainly eroded away.

A mandible sketch in the sand I did to explain what we were looking at.


I routinely make sketches from memory in the sand - why not? It's the world's biggest white board! And it usually erases itself at the next high tide.

 Wayne helping Sarah off the beach while I was being super helpful and taking pictures...

We drove down to another spot and took a trail down one of the larger gullies - one other spot afforded a trail down nearly a mile closer to the best fossil-bearing spot, but unfortunately this more conveniently located trail is typically overgrown with poison oak. We chatted about the Putah tuff-correlative ash exposed in this gully, here nearly a meter thick - Jorge explained that while the Ishi tuff-correlative ash further up-section within the Purisima Formation is several meters thick, the actual ash layer near the volcanic vent itself is only around 10-15 cm thick. This means that the ash layer has been dramatically exaggerated through sedimentation - this ash was likely deposited inland and redeposited as a thick layer on the continental shelf by a river mouth on the periphery of the Purisima depositional basin, perhaps the ancient opening of the San Joaquin embayment (the extinct "Priest Valley Strait", which used to connect the marine San Joaquin embayment to the formerly marine Salinas River Valley) to the southeast. 

 

A great exposure of the ash bed correlative with the 3.35 Ma Putah Tuff - white ash-filled burrows extend downwards into the brownish gray Purisima sandstone, and brownish-gray sediment infills burrows extending into the top of the ash. The ash bed iself is about 1-1.5 meters thick and cross-bedded - hummocky cross-beds, I believe - indicating rapid sedimentation, too rapid for the sediment to be bioturbated by burrowers.


 An impression of a large scallop, likely
Patinopecten healyi - a Pliocene index fossil in California.


 A fractured boulder with a bed of slipper snails (
Crepidula princeps) - these gastropods lived in large clusters and are hermaphroditic, often forming small patch reefs. Shell beds like this can be a few meters across. In this case, the shell has been dissolved away - common in this section. However, these shells are also frequently bored into owing to their non-burrowing ecology - the little rice grain sized blobs are the infill of boring sponges (Clionidae), the trace fossil Entobia.

On our way to the best collecting area we encountered our first rockfall - a brand new one that must have occurred in the past five months since I had not seen it on my December visit. Some of the boulders exposed shell bed layers that had in the past (like, in 2006) produced some rare teeth (e.g. a sawshark rostral tooth) and a tympanic bulla of a true porpoise (Phocoenidae). I saw some interesting crustacean parts and then immediately found two different tympanic bullae - one appeared to be from a phocoenid or perhaps a delphinid (oceanic dolphin), and the other one was very clearly from Parapontoporia, the "river dolphin" from the Purisima Formation that is most closely related to the recently extinct Chinese river dolphin (Lipotes vexillifer). I was quite pleased - in just a few minutes, I raised the number of dolphin bullae from three to five, nearly doubling it! These were somewhat more common than the more taxonomically informative periotic bones - only two of which had ever been found: one I found in 2010 or 2011, and another that Chris Pirrone had found with me. 


Walking down along the cliffs to the best collecting area.


 Formidable cliffs.

We visited the two most promising localities, and unfortunately neither produced much: I was hoping for a couple of shark teeth or marine mammal teeth, maybe a periotic bone. Nada; just a few bone fragments. The cliffs here didn't look too different than in December, and even more erosion would likely be needed. I had hoped that by bringing Sarah along she'd find some teeny tiny little skate or stingray teeth from one micro-tooth bearing shell bed, but she had been quite cold and tired and wasn't in her prime. I managed to do somewhat better than past visits and found three or four 1-2mm wide skate (Raja) and/or stingray (Dasyatis) teeth - so that was nice.

 Sarah smiling despite being quite chilly on the walk down.

Yours truly with a tiny little fur seal mandible! I'll post more photos once it's prepared. Easily the find of the day.

Luck finally changed when we made it down to the last stop, which was a sea cave that has grown considerably since I first found bones in it in 2005. In December I had collected an unusual looking bone and upon pulling it out, realized it was the rostrum of a dolphin, probably Parapontoporia - I did not have the time or equipment to collect the rest of the skull, so I hoped it would survive the winter storms that were brewing on the horizon. As I entered the cave and saw how low the sand was and how high the ceiling was, my heart sank a bit. No trace of the dolphin could be located after about 20 minutes of desperate searching. Few newly exposed bits of bone looked enticing. I finally spotted a tiny bone about 8 feet above the sand that looked intriguing. After a few failed attempts of scaling the wall of the cave I almost gave up, but on the last try I used some footholds I had carved in years ago and scaled the wall and traversed over to it - I stuck my chisel in and gave it a couple of taps and saw that it was coming loose. I put my hammer back on my belt, and while precariously balanced, I used one hand to cradle the specimen and the other hand to delicately push and leverage the chisel into the rock and the little bone tilted over gently into my hand. I briefly caught glinting of enamel; before I thought I might lose my balance, I pocketed my chisel and looked more closely - I saw two teeth. "Holy shit I found something great" I exclaimed as I hopped down to the sand. I held it up and realized I was staring at a tiny little fragment of mandible with two teeth in place - teeth clearly identifiable as pinniped - and probably a fur seal. My luck had finally turned around! "I could leave right now and be happy with today!" Sarah, who was happy for me but quite chilled, snidely remarked "let's leave now then." I showed the find to the rest of the group and explained how rare pinniped fossils are, and that I had been desperately looking for such a fossil from this site for nearly 20 years. This specimen now marks the fourth pinniped mandible from the Purisima I've collected in the past 16 months! We walked a bit further to take a look at some ash beds, and then headed back up towards the trail.


 

When we got back to the gully we were going to walk back up, we saw a sea lion on the beach - I thought at first that it was a very fresh carcass. Then, I saw it breathe and its nostrils flared. It seemed to be a 2-3 year old male, and not aware of us - assuming it was a carcass, we got within 15 feet - until we realized it was unconscious. It was not starving (no ribs/scapulae/knees protruding) and had no obvious lesions or bite marks on it - I assumed that it was sick. Wayne Thompson observed that it was shallowly breathing - indeed, breathing quite slowly, and called the Marine Mammal Center. Normally, when you approach a sea lion, they open their eyes and will shift their weight while they're judging whether or not you're a threat - and even if they're not aware of you, they often snort and scratch themselves while asleep. 

The tympanic bulla of a monodontid as first located after Sarah slipped.


The periotic of the monodontid, found about a meter away. The white weathered bit - the posterior process - was all that was exposed.


A bit of chiseling...


 ...and voila! Happy whaleontologist with a new record for the locality.

We started heading back up the gully, and I tried to help Sarah up a small ledge out of some rancid water (water backed up behind a sandbar in the gully that was filled with decomposing Velella velella, by-the-wind-sailors). While I was pulling her up, one of her feet slipped and she banged up her knee pretty good - she cussed for a couple minutes but was otherwise OK. While she was rubbing her knee and resting for a moment, I noticed what I first thought was the mold of a mollusk shell - and then realized it was actually a pretty chunky tympanic bulla from an odontocete! My third one for the day. As I chiseled it out I noted that it's a bit larger than the most common tympanic bullae from the Purisima, meaning that it's not Parapontoporia or from a phocoenid porpoise - or a small delphinid. I thought at first it might be from a globicephaline, like a pilot whale, which I already documented from this locality a decade ago. While I was chatting with the others and wrapped the specimen up, I saw a little 1 cm wide bit of bone sticking out - it had some distinctive little spurs and longitudinal grooves that I immediately recognized as part of the posterior process of another earbone, about a meter away. At first I thought this was likely the other tympanic bulla. As I looked closer, I remarked "No %%%-ing way" and began to carefully chisel: it was a periotic! The periotic is the more informative of the two earbones. After a minute I confirmed that this was a large odontocete, possibly a pilot whale or maybe from a beluga (Monodontidae). As the periotic popped out without fracturing, I saw the dorsal side - confirming it to be from a monodontid! With this associated pair of earbones, I had doubled the sample of odontocete bullae, and increased the sample of odontocete periotics from this locality from two to three. Further, this is the first confirmed monodontid from this locality in the Purisima Formation. Needless to say, I was elated! So, I thanked my dear wife for falling and getting a bruised up knee for a couple days: I think I'll have to list her as a co-finder for her role in the find.


 As everywhere else along the Pacific coast, there was no shortage of by-the-wind-sailors (Velella velella) washing up along the beach here.

A minute later, Wayne indicated that he had found some fish bones - he brought over a chunk of rock he had chiseled out with a couple of vertebrae. "And there's more in the rock!", he said. I came over there, and sure enough, there were a few additional bones including something that looked symmetrical - perhaps a braincase. The vertebrae were about 2 cm long or so. Wayne and I collected the last few bits and bagged them up. At this point, Sarah noticed that the sea lion had woken up and was standing up at attention, looking at us with mild concern. It was getting a bit late in the afternoon and Wayne parted ways - the rest of us headed a bit further north to get to the localities we had wanted to check earlier in the morning, but were barred by waves. Sarah was a bit cold and tired (she informed us we had hit five miles of walking a bit earlier) and opted to stay in the car while Jorge, Marsha, and I walked south. Jorge and Marsha pointed out different ash beds they had sampled, and I checked the last few localities.


 The Herpetocetus skull, still in its faithful concretion, waiting patiently for us to excavate it on a followup visit later in the summer.


 A partial palatoquadrate cartilage of a skate,
Raja binoculata. This is the lower jaw, and is composed of calcified cartilage - often fibrous but with a layer of prismatic (tesselated) cartilage.


A partial rostrum of a medium-sized cetacean - most likely a small mysticete, but something about it looks odd to me.

I confirmed that the Herpetocetus skull I found last year survived all of the winter erosion, which was a relief! And further yet, not covered in sand. A few feet away I found a nice palatoquadrate cartilage of a skate (Raja). A few minutes later I found an unusual bone that had not been exposed in December, which came out in a few pieces. A bit further on down the beach I pointed out a minke whale-sized mandible which I had wanted to excavate, but half of it was removed by a fossil poacher. I then located what appeared to be a relatively flat looking rostrum from a cetacean - which is either from a dwarf whale like Herpetocetus or perhaps from a monodontid. A follow up visit is necessary to excavate this.

A greenish chunk of metaconglomerate, likely from the Franciscan complex - the rocks that make up the hills in San Francisco, much of the east bay, Angel Island, Alcatraz, and the Marin Headlands.

 A cluster of sand dollars, probably Merriamaster.


 A much younger ecentric sand dollar (
Dendraster excentricus) with a huge red striped acorn barnacle (Paraconcavus pacificus) stuck to it, with several additional barnacles. A common occurrence along this section of the coast!


A closeup of some slipper snail colonies (Crepidula princeps), along with a large scallop.


The entire shell bed - a concretion about 1.5x2 meters in size, 99% of the fossils belonging to a single species. This would be a 'community bed' according to Richard Norris' (1986) research on the taphonomy of the Purisima Formation.